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Wall, and extrudes to encapsulate the seed. The extruded Arabidopsis seed mucilage has at least two distinct layers, nonadherent and adherent (Western et al., 2000; Macquet et al., 2007a). The outermost layer (nonadherent layer) is amorphous in appearance, composed primarily of pectin and, as its name suggests, easily separated from the seed by gentle shaking. The layer of mucilage adjacent to the

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